My brother Tim's 3C had a similar problem a few years back. It suddenly started running as rich as buggery on the centre cylinder, as anyone who rode behind him on the way to the last Laverda rally at Bellingen could attest. They were choking on the black smoke coming out of his pipes! When he got back home we threw another set of carbs on the bike (they were off my Jota after I'd fitted a set of Mikunis). That fixed the problem, so we left those carbs on the bike. His old carbs were chucked in a box and haven't been looked at to this day, so we never did get any closer to a proper diagnosis other than confirming it was a carby issue. Not much help eh?
As both yours and Tim's rich running problem had a sudden onset, I'd be looking for something that could fail suddenly. There are a couple of things that come to mind ...
Check that the rubber tip on the float needle hasn't fallen off, and that the float valve is still screwed into the carb body with an good gasket.
Also check that the float still does what the name says (floats) and the tab that closes the valve hasn't broken off (seen that once - must have been adjusted once too often) .
I see that you've eliminated the choke plungers as a problem by swapping them around (not actually a choke, but we all call it that). But have a look at the choke valve seat on the middle carb. It's unlikely to be a problem (maybe a chunk fallen out of the edge of the seat or a piece of crud in there stopping the thing from closing) but worth casting your eye on it just to be sure.
BTW, if anyone has a problem with the choke valves not sealing very well, check that the rubber seal in the end of the brass piston isn't worn out. If the circular groove where it sits on top of the valve opening gets too deep, sometimes the plunger can hang on the edge of the groove and not seal. Rather than shelling out a handful of money for new choke plungers, the first thing to try is to pick the rubber seal out of the end and flip it over to expose a fresh surface. If you've already done that and worn out both sides, you can't buy replacement rubbers but you can make them. Get hold of some 3mm fuel resistant rubber sheet and punch them out with a hole punch - 6mm dia I think (memory is a bit hazy on that, so check the punch diameter against the old rubbers). 3mm (1/8") thick x 100mm square Viton sheet is available on eBay for about $8. That's about the cost of a single choke plunger, so if you have a bunch to do it's worth doing. If you regularly work on Dellorto carbs, it'll save quite a bit in the long run.